Ignore the top speed comments. Assuming you're talking about a CJ/YJ/TJ (Wrangler) then 4.56 gearing is roughly appropriate for a 6 cyl engine with 35" tires. If you're talking about a 4 cyl with 35" tires, something like 4.88 would be more appropriate. The gears serve to compensate for the overly large tires. A "stock" TJ Wrangler would have 3.06 or 3.73 gears with the Inline 6 cyl, or 4.11 gears with the 4 cyl. That would be with a stock 30" tire. (technically, there are 2 stock tire sizes, but that's not important right now)
Basically it means that 4.56 revolutions of the engine = 1 revolution of the wheels. So if your gearing was 2:1, then at 1000 rpm your wheels would be turning at 500 rpm. Make sense? You can find the diameter of the wheel, and use that to get actual speeds for given revolutions. Or go to some Jeep websites (I know there is one at
www.dirtroad.com) and you'll find a gear ratio / tire size calculator.
Any jeep questions, feel free to shoot me an email. (not PM, don't check those as often)